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The grass genera of the world

L. Watson, T.D. Macfarlane, and M.J. Dallwitz

Nianhochloa H.N. Nguyen & V.T. Tran

Type species: N. bidoupensis H.N. Nguyen & V.T. Tran.

Habit, vegetative morphology. Perennial; probably caespitose. The flowering culms leafy. Culms 300–800; woody and persistent; to 4 cm in diameter; cylindrical; scrambling or scandent (?); branched above. Primary branches derived from a single bud, 3–4 (the branches subequal to the internode); clumped (with subequal branches). The branching dendroid. Culm nodes slightly swollen. Culm leaf sheaths present; deciduous. Culm leaves with conspicuous blades. Culm leaf blades triangular. Culm internodes 20–22 cm long, hollow. Rhizomes short, pachymorph. Leaves not basally aggregated; non-auriculate. Leaf blades oblong- ovate; broad; 14–16 cm long, 25–28 mm wide; not cordate, not sagittate (broadly rounded); pseudopetiolate; with 10–12 secondary veins; disarticulating from the sheaths. Ligule a fringed membrane; 1 mm long.

Reproductive organization. Plants bisexual, all with bisexual spikelets; with hermaphrodite florets. The spikelets hermaphrodite.

Inflorescence. Inflorescence indeterminate; with pseudospikelets; (synflorescence) bractiferous, clustered at the nodes, in untidy tufts, lax, with axillary buds at the bases of the spikelets, prophyllate below lateral spikelets, leafless between clusters.

Female-fertile spikelets. Spikelets unconventional; 10–20 mm long; lanceolate; compressed laterally; disarticulating above the glumes; disarticulating between the florets; with conventional internode spacings (internodes 2–3 mm long, glabrous). Rachilla prolonged beyond the uppermost female-fertile floret.

Glumes one per spikelet to several (1, 2 or 3 empty glumes, preceded by one or two gemmiferous bracts); shorter than the spikelets; (at least the the upper) hairy (pilose); (the upper) pointed (acuminate and mucronate); thinner than the fertile lemma, similar. Spikelets with incomplete florets. The incomplete florets distal to the female-fertile florets. The distal incomplete florets 1; merely underdeveloped; awnless. Spikelets without proximal incomplete florets.

Female-fertile florets (1–)2(–3). Lemmas ovate; not convolute; decidedly firmer than the glumes; 7–9 mm long, entire; pointed, or blunt (?); mucronate; hairless; glabrous; non-carinate; 9–10 nerved. Palea present; relatively long (as long as the lemma); hyaline thinner than the lemma; not indurated; several nerved (9–10); 2-keeled. Palea back glabrous. Lodicules present (2–3 mm long); 3; basaly fleshy and protruding, apically transparent and membranous; protruding, glabrous. Stamens 6; with free filaments. Anthers 4–5 mm long. Ovary densely apically hairy; without a conspicuous apical appendage (cf. Nguyen & Tran illustration). Styles short. Stigmas plumose, 3.

Fruit, embryo and seedling. Fruit medium sized (5 mm long); ellipsoid (apically rostrate); compressed dorsiventrally, or not noticeably compressed (compressed on one side). Embryo large.

Classification. Watson & Dallwitz (1994): not described separately. Soreng et al. (2015): Bambusoideae; Bambusodae; Bambuseae; Bambusinae. 1 species (N. bidoupensis).

Distribution, phytogeography, ecology. Bidoup Mountain, S. Vietnam.

References, etc. Morphological/taxonomic: Bguyen, H.N. and Tran, V.T. (2012): Adansonia 34, 257–264; Clayton et al. (2016), Grassbase.

Special comments. Anatomical data wanting.


We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, distributions of character states within any set of taxa, geographical distribution, and classifications. See also Guidelines for using data taken from Web publications.


Cite this publication as: ‘Watson, L., Macfarlane, T.D., and Dallwitz, M.J. 1992 onwards. The grass genera of the world: descriptions, illustrations, identification, and information retrieval; including synonyms, morphology, anatomy, physiology, phytochemistry, cytology, classification, pathogens, world and local distribution, and references. Version: 13th November 2017. delta-intkey.com/grass’.

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