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The grass genera of the world

L. Watson, T.D. Macfarlane, and M.J. Dallwitz

Ampelodesmos Link

Habit, vegetative morphology. Robust perennial; rhizomatous. Culms 60–350 cm high; herbaceous. Culm internodes solid. Leaves non-auriculate. Leaf blades harsh; narrow; flat, or rolled; without cross venation. Ligule a fringed membrane; not truncate; 6–12 mm long.

Reproductive organization. Plants bisexual, all with bisexual spikelets; with hermaphrodite florets.

Inflorescence. Inflorescence paniculate; open; espatheate; not comprising ‘partial inflorescences’ and foliar organs. Spikelet-bearing axes persistent. Spikelets not secund; pedicellate.

Female-fertile spikelets. Spikelets 10–15 mm long; compressed laterally; disarticulating above the glumes; disarticulating between the florets. Rachilla prolonged beyond the uppermost female-fertile floret; hairy (villous). Hairy callus present.

Glumes two; more or less equal; shorter than the spikelets; shorter than the adjacent lemmas; pointed (acuminate); awnless; carinate; similar (firmly membranous). Lower glume 3–5 nerved. Upper glume 3–5 nerved. Spikelets with female-fertile florets only, or with incomplete florets. The incomplete florets distal to the female-fertile florets.

Female-fertile florets 2–6 (?). Lemmas decidedly firmer than the glumes (leathery); not becoming indurated; incised; 2 lobed; not deeply cleft (shortly bidentate); mucronate, or awned. Awns when present, 1; from a sinus, or apical; non-geniculate; much shorter than the body of the lemma. Lemmas hairy (on the lower half); carinate to non-carinate; 5–7 nerved. Palea present; relatively long; apically notched (bidentate); awnless, without apical setae; 2-nerved; 2-keeled. Lodicules present; 3; free; membranous; ciliate (on the margins); not toothed; not or scarcely vascularized. Stamens 3. Anthers not penicillate. Ovary apically hairy; with a conspicuous apical appendage. Styles free to their bases. Stigmas 2; white.

Fruit, embryo and seedling. Fruit medium sized (about 7 mm long); not grooved; with hairs confined to a terminal tuft. Hilum long-linear. Embryo small. Endosperm hard. Embryo with an epiblast; without a scutellar tail; with a negligible mesocotyl internode. Embryonic leaf margins meeting.

Seedling with a tight coleoptile. First seedling leaf with a well-developed lamina. The lamina narrow (l/b ratio 70); curved; 7 veined.

Abaxial leaf blade epidermis. Costal/intercostal zonation lacking. Papillae absent. Long-cells similar in shape costally and intercostally; of similar wall thickness costally and intercostally. Mid-intercostal long-cells rectangular; having markedly sinuous walls. Microhairs absent. Stomata absent or very rare; 28.5–30 microns long. Intercostal short-cells common; in cork/silica-cell pairs. Intercostal silica bodies elliptic. Costal short-cells predominantly paired. Costal silica bodies rounded (mostly, elliptical), or tall-and-narrow.

Transverse section of leaf blade, physiology. C3; XyMS+. Mesophyll with non-radiate chlorenchyma. Leaf blade with distinct, prominent adaxial ribs (flat-topped ribs); with the ribs very irregular in sizes. Midrib not readily distinguishable; with one bundle only. Bulliforms not present in discrete, regular adaxial groups; in the furrows, in ill-defined groups of irregularly sized cells cf. Ammophila. All the vascular bundles accompanied by sclerenchyma. Sclerenchyma not all bundle-associated. The ‘extra’ sclerenchyma in a continuous abaxial layer.

Cytology. Chromosome base number, x = 12 (chromosomes small). 2n = 48 and 96. 4 and 8 ploid.

Classification. Watson & Dallwitz (1994): Stipoideae; Ampelodesmeae. Soreng et al. (2015): Pooideae; Ampelodesmeae. 2 species (A. ampelodesmos, A. mauritanicus).

Distribution, phytogeography, ecology. Mediterranean.

Xerophytic (mainly coastal).

Economic aspects. A component of Esparto grass, used for papermaking.

References, etc. Morphological/taxonomic: Decker 1964b; Macfarlane and Watson 1980. Leaf anatomical: Metcalfe 1960; studied by us - A. mauritanicus (Poir.) Dur. & Schinz.

Special comments. An isolated monotypic genus, of uncertain taxonomic affinities in spite of the good descriptive data. A prime candidate for comparative DNA studies. Illustrations. • A. mauritanicus, as Arundo bicolor: Desfontaines (1798), Fl. Atlantica 1. • A. mauritanicus, as Deyeuxia arundinacea: P. Beauv. (1812). • A. mauritanicus, abaxial epidermis of leaf blade: this project. • A. mauritanicus, transverse section of leaf blade: this project

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, distributions of character states within any set of taxa, geographical distribution, and classifications. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Watson, L., Macfarlane, T.D., and Dallwitz, M.J. 1992 onwards. The grass genera of the world: descriptions, illustrations, identification, and information retrieval; including synonyms, morphology, anatomy, physiology, phytochemistry, cytology, classification, pathogens, world and local distribution, and references. Version: 11th December 2017.’.