Festuca of North America


S.G. Aiken, M.J. Dallwitz, C.L. McJannet, and L.L. Consaul


Abel, D.J., and Williams, W.T. 1985. A re-examination of four classification fusion strategies. Comput. J. 28: 439–443.

Abrams, L. 1923. An illustrated flora of the Pacific States, Stanford University Press, Washington, Oregon and California, USA. Vol. 1 of 4.

Aiken, S.G. 1993. Festuca. In The Jepson Manual: higher plants of California. ed. J.C. Hickman. University of California Press, Berkeley, Los Angeles. 1259–1262.

Aiken, S.G., and Consaul, L.L. 1995. Leaf cross sections and phytogeography: a potent combination for identifying members of Festuca subgg. Festuca and Leucopoa (Poaceae) occurring in North America. Amer. J. Bot. 82: 1287–1299.

Aiken, S.G., Consaul, L.L., Davis, J.I., and Manos, P.S. 1993. Systematic inferences from variation in isozyme profiles of arctic and alpine cespitose Festuca (Poaceae). Amer. J. Bot. 80: 76–82.

Aiken, S.G., Consaul, L.L., and Lefkovitch, L.P. 1995. Festuca edlundiae (Poaceae), a High Arctic, new species compared enzymatically and morphologically with similar Festuca species. Sys Bot. 20: 374–392.

Aiken, S.G., Consaul, L.L., Spidle, A., and May, B. 1994. Allozyme and morphological observations on Festuca hyperborea compared with F. baffinensis and F. brachyphylla (Poaceae) from the Canadian Arctic. Nord. J. Bot. 14: 137–143.

Aiken, S.G., and Dallwitz, M.J. 1991. Festuca of North America - Interactive identification and information retrieval. Taxacom: Flora Online, 26. Data bank on floppy disc.

Aiken, S. G., Dallwitz, M. J., McJannet, C. L., and Consaul, L. L. (1997). Biodiversity among Festuca (Poaceae) in North America: diagnostic evidence from DELTA and clustering programs, and an INTKEY package for interactive, illustrated identification and information retrieval. Can. J. Bot. 75, 1527–1555.

Aiken, S.G., and Darbyshire, S.J. 1984. The morphology of a vegetatively proliferating inflorescence of Kentucky Bluegrass, Poa pratensis. Can. Field-Naturalist, 98: 249–251.

Aiken, S.G., and Darbyshire, S.J. 1990. Fescue grasses of Canada. Agriculture Canada Research Branch, publication 1844/E.

Aiken, S.G., Darbyshire, S.J., and Lefkovitch, L.P. 1985. Restricted taxonomic value of leaf section in Canadian narrow-leaved Festuca (Poaceae). Can. J. Bot. 63: 995–1007.

Aiken, S.G. and Fedak, G. 1992. Chromosome numbers in North American Festuca (Poaceae). Can. J. Bot. 70: 1940–1944.

Aiken, S.G., and Gardiner, S.E. 1991. SDS-PAGE of seed proteins in Festuca (Poaceae): taxonomic implications. Can. J. Bot. 69: 1425–1432.

Aiken, S.G., Gardiner, S.E., and Forde, M.B. 1992. Taxonomic implications of SDS-PAGE analyses of seed proteins in North American taxa of subgenus Festuca (Poaceae). Biochemical Systematics and Ecol. 20: 615–629.

Aiken, S.G. and Lefkovitch, L.P. 1984. The taxonomic value of using epidermal characteristics in the Canadian rough fescue complex (Festuca altaica, F. campestris, F. hallii, "F. scabrella"). Can. J. Bot. 62: 1864–1870.

Aiken, S.G., and Lefkovitch, L.P. 1990. Arctagrostis (Poaceae, Tribe Pooideae) in North America and Greenland. Can. J. Bot. 68: 2422–2432.

Aiken, S.G., and Lefkovitch, L.P. 1993. On the separation of two species of Festuca subg. Obtusae (Poaceae). Taxon, 42: 323–337.

Aiken, S.G., Lefkovitch, L.P., Darbyshire, S.J., and Armstrong, K.C. l987. Vegetative proliferation in inflorescences of red fescue (Festuca rubra L. s.l. Poaceae). Can. J. Bot. 66: 1–10.

Ainscough, M.M., Barker, C.M., and Stace, C.A. 1986. Natural hybrids between Festuca and species of Vulpia. Watsonia, 16: 143–151.

Albee, B.J., Shultz, L.M., and Goodrich, S. 1988. Atlas of the vascular plants of Utah. Utah Museum of Natural History. Occasional Publications 7. Salt Lake City, Utah, USA.

Aldon, E.F., and Barsted, J.F. 1987. Escudilla Mountain Research Natural area: a study of an undisturbed montane grassland in Arizona. Natural Areas Journal, 7: 107–117.

Alexeev, E.B. 1985. Festuca L. (Poaceae) in Alaska and Canada. Novst. Sist. Vyssh. Rast. 22: 5–35 (in Russian).

Alexeev, E.B. 1984. Viviparous fescues (Festuca L., Gramineae) of South America. Byull. Mosk. O-Va Ispyt. Prir. Otd. Biol. 89: 111–117 (in Russian).

Alexeev, E.B. 1983. Two new narrow-leaved species of the genus Festuca (Gramineae) from North America. Byull. Mosk. O-Va Ispyt. Prir. Otd. Biol. 88: 103–104.

Alexeev, E.B. 1982. New and little known Festuca of North America. Byull. Mosk. O-Va Ispyt. Prir. Otd. Biol. 89: 111–117.

Alexeev, E.B. 1975. Generis Festuca L. species angustifoliae partis Europaeae U.S.S.R. Novst. Sist. Vyssh. Rast. 17: 42–53 (in Russian).

Alexeev, E.B. 1972. The significance of degree of integration of leaf vaginae of vegetative shoots in the systematism of Fescues of the group Festuca ovina L. s.l. Vestn. Mosk. Univ. Ser. 16 Biol. 5: 48–51 (in Russian).

Allred, K.W. 1993. A field guide to the grasses of New Mexico. Agricultural Experiment Station, New Mexico University, New Mexico, USA.

Angelov, G.B. 1993. A biosystematic study on species of genus Festuca L. 5. An electrophoretic analysis of the enzymes glucose-6-phosphate dehydrogenase, 6-phosphogluconate dehydrogenase and superoxide dismutase. Fitologija, 44: 47–56.

Angelov, G.B. 1992. Biosystematic study of species from genus Festuca L. 4. Electrophoretic analysis of the enzymes peroxidase, acid phosphatase and catalase. Fitologija, 43: 58–68.

Argus, G.W., and Aiken, S.G. 1987. Noteworthy collections. New Mexico. Festuca minutiflora Rydb. Madroño, 34: 267.

Atkinson, W.D., and Gammerman, A. 1987. An application of expert systems technology to biological identification. Taxon, 36: 705–714.

Auquier, P. 1973. Festuca. In Nouvelle flore de la Belgique, du Grand-Duché de Luxembourg, du Nord de la France et des régions voisines. eds. J.-E. de Langhe et al. Jard. Bot. Natl. Belg., Brussels, Belgium.

Bardoux, S. 1971. Sur l'anatomie de la feuille de Festuca arundinacea Schreb., Festuca pratensis Huds., Lolium multiflorum Lam. et leurs hybrides. Bull. Soc. Vaudoise Sci. Nat. 71: 15–22.

Barkley, T.M. ed. 1977. Atlas of the flora of the Great Plains. Iowa State University Press, Ames, Iowa, USA.

Baun, E.L. 1950. Deciduous forests of Eastern North America. Blakiston Co. Toronto, Canada.

Beddows, A.R. 1931. Seed setting and flowering in various grasses. Welsh Pl. Breed. St. Bull. Ser. H. Nr. 12: 5–99.

Blood, D.A. 1966. The Festuca scabrella association in Riding Mountain national park, Manitoba. Can. Field-Naturalist, 80: 24–32.

Boivin, B. 1981. Flora of the Prairie Provinces. A handbook to the flora of the provinces of Manitoba, Saskatchewan, and Alberta. Part 5. Gramineae. Provancheria, 12.

Boivin, B. 1967. Enumération des plantes du Canada. 6 Monopsides. Naturaliste Can. (Qué.), 94: 471–528.

Borrill, M., Kirby, M., and Morgan, G. 1980. Studies in Festuca. 12. Morphology, distribution and cytogenetics of F. donax, F. scariosa and their hybrids, and the evolutionary significance of their fertile amphiploid derivative. New Phytol. 86: 423–439.

Borrill, M., Kirby, M., and Morgan, W.G. 1977. Studies in Festuca. 11. Interrelationships of some putative diploid ancestors of the polyploid broad-leaves fescues. New Phytol. 78: 661–674.

Borrill, M., Tyler, B.F., and Morgan, W.G. 1976. Studies in Festuca. 7. Chromosome atlas (Part 2) An appraisal of chromosome race distribution and ecology, including F. pratensis var. apennina (De Not.) Hack. tetraploid. Cytologia, 41: 219–236.

Borrill, M. 1972. Studies in Festuca. 3. The contribution of F. scariosa to the evolution of polyploids in sections Bovinae and Scariosae. New. Phytol. 71: 523–532.

Borrill, M., Tyler, B., and Lloyd-Jones, M. 1971. Studies in Festuca. 1. A chromosome atlas of Bovinae and Scariosae. Cytologia, 36: 1–14.

Bowden, W.M. 1960. Chromosome notes and taxonomic notes on northern grasses. 2. Tribe Festuceae. Can. J. Bot. 38: 541–557.

Brayshaw, T.C. 1970. The dry forests of British Columbia. Syesis, 3: 17–43.

Brayshaw, T.C. 1965. The dry forests of southern British Columbia. Ecol. W. N. Amer. 1: 65–75.

Bruhl, J., Watson, L., and Dallwitz, M.J. 1992. Genera of Cyperaceae: interactive identification and information retrieval. Taxon, 41: 225–233.

Buckner, R.C., and Jackson, J.A. 1982. Update on tall fescue varieties and their uses. Dairy Research Report. Kentucky Agricultural Experiment Station. 264: 22.

Buckner, R.C., and Bush, L.P. 1979. eds. Tall fescue. Amer. Soc. Agronomy, 20. Crop Sci. Soc. Amer. and Soil Sci Soc. Amer. Inc., publishers. Madison. Wisconsin, USA.

Bulinska-Radomska, Z., and Lester, R.N. 1988. Integeneric relationships of Lolium, Festuca, and Vulpia (Poaceae) and their phylogeny. Pl. Syst. Evol. 159: 217–227.

Bulinska-Radomska, Z., and Lester, R.N. 1986. Phylogeny of chromosome races of Festuca arundinacea and F. mairei (Poaceae) as indicated by seed protein electrophoresis. Pl. Syst. Evol. 152: 153–166.

Bulinska-Radomska, Z., and Lester, R.N. 1986. Species relationships in Festuca (Sect. Ovinae, Poaceae). Pl. Syst. Evol. 154: 175–182.

Burner, D.M., Balasko, J.A., and O'Brien, P.M. 1988. Attributes of tall fescue germplasm of diverse geographic origin. Crop Sci. 28: 459–462.

Burr, E. J. 1970. Cluster sorting with mixed character types. 2. Fusion strategies. Austral. Comput. J. 2: 98–103.

Butkute, B.L., and Konarev, A.V. 1982. Studies of the seed proteins in the genera Lolium and Festuca (Poaceae) in connection with their phylogeny. Bot. Zhur. 67: 812–819.

Butkute, B.L., and Konarev, A.V. 1981. An immunochemical study of the seed proteins of Lolium in connection with the phylogeny of this genus. Bot. Zhur. (Lenn.) 65: 1453–1458.

Chandrasekharan, P., Lewis, E.J., and Borrill, M. 1972. Studies in Festuca. 2. Fertility relationships between species of sections Bovinae and Scariosae, and their affinities with Lolium. Genetica, 43: 375–386.

Chase, A., and Niles, C.D. 1962. Index to grass species. Vol. 2. G.K. Hall. Boston, Massachusetts, USA

Clarke, J., Chandrasekharan, P., and Thomas, H. 1976. Studies in Festuca. 9. Cytological studies of Festuca pratensis var. apennina (De Not.) Hack. (2n=28). Z. Pflanzenzüchtg. 77: 205–214.

Cohen, C.J., Chilcote, D.O., and Frakes, R.V. 1982. Leaf anatomy and stomatal characteristics of four tall fescue Festuca arundinacea selections differing in forage yield. Crop Sci. 22: 704–708.

Cohen, C.J., Chilcote, D.O., and Frakes, R.V. 1982. Gas exchange and leaf area characteristics of four tall fescue Festuca arundinacea selections differing in forage yield. Crop Sci. 22: 709–711.

Connor, H.E. 1968. Interspecific hybrids in hexaploid New Zealand Festuca. New Zealand J. Bot. 6: 295–308.

Connor, H.E. 1960. Variations in leaf anatomy in Festuca novae-zelandia (Hack.) Cockayne and F. matthewsii (Hack.) Cheeseman. New Zealand J. Science, 3: 468–509.

Consaul, L.L., and Aiken, S.G. 1993. Limited taxonomic value of palea intercostal characteristics in North American Festuca (Poaceae). Can. J. Bot. 71: 1651–1659.

Correll, D.S., and Johnson, M.C. 1970. Manual of the Vascular Plants of Texas. Texas Research Foundation, Renner, Texas, USA.

Cronquist, A., Holmgren, A.H., Holmgren, N.H., and Reveal, J.L. 1977. Intermountain flora. Vascular Plants of Intermountain West, U.S.A. Vol. 6. Columbia Univ. Press, New York, New York, USA.

Crowder, L.V. 1956. Morphological and cytological studies in tall fescue (Festuca arundinacea Schreb.) and meadow fescue (F. elatior L.). Bot. Gaz. 117: 214–223.

Dallwitz, M.J. 1980. A general system for coding taxonomic descriptions. Taxon, 29: 41–46.

Dallwitz, M J., Paine, T.A., and Zurcher, E.J. 1993 onwards. User’s guide to the DELTA system: a general system for processing taxonomic descriptions. 4th edition. http://delta-intkey.com/

Dallwitz, M.J., Paine, T.A., and Zurcher, E.J. 1995 onwards. User’s guide to Intkey: a program for interactive identification and information retrieval. 1st edition. http://delta-intkey.com/

Dallwitz, M. J., Paine, T. A. and Zurcher, E. J. 2000 onwards. Principles of interactive keys. http://delta-intkey.com/

Darbyshire, S.J. 1993. Realignment of Festuca subgenus Schedonorus with the genus Lolium (Poaceae). Novon, 3: 239–243.

Darbyshire, S.J., and Warwick, S.I. 1992. Phylogeny of North American Festuca (Poaceae) and related genera using chloroplast DNA restriction site variation. Can. J. Bot. 70: 2415–2429.

Dayton, W.A. ed. 1937. Range plant handbook. U.S. Forest Service, Washington, D.C. U.S. Dept. of Commerce/ National Bureau of Standards/ Institute of Applied Technology.

Dore, W.G., and McNeill, J. 1980. Grasses of Ontario. Agric. Can. Monogr. 26.

Drobow, V. 1915. Predstavitieli sekcii Ovinae Fr. roda Festuca L. v Jakutskoj oblasti. Trav. Mus. Bot. Acad. Imp. Sci. Petrograd, 14: 147–171.

Druce, G.A. 1915. Plant notes etc. for 1914. Rep. Bot. Soc. Exch. Club Br. Isl. 4: 1–48.

Dubé, M., and Morisset, P. 1995. Novelles observations cytologiques chez Festuca rubra sensu lato (Poaceae) dans l'est du Canada. Can. J. Bot. 73: 1289–1294.

Dubé, M., and Morisset, P. 1987. Morphological and leaf anatomical variation in Festuca rubra sensu lato (Poaceae) from eastern Quebec. Can. J. Bot. 65: 1065–1077.

Dubé, M., Morisset, P., and Murdock, J. 1985. Races chromosomiques chez Festuca rubra sensu lato (Poaceae) dans l'est du Québec. Can. J. Bot. 63: 227–231.

Dubé, M., and Morisset, P. 1983. Addition de Festuca gigantea (L.) Vill. (Poaceae) à la flore du Canada. Naturaliste Can. (Qué.), 110: 213–215.

Dulepova, B.I. 1986. Ecological and phytocenotic features of narrow-leaved fescues in the Dauria forest-steppe. Ecol. 26: 72–73.

Duyvendak, R., Luesink, B., Vos, H. 1981. Delimitation of taxa and cultivars of red fescue (Festuca rubra L. sensu lato) Rasen, 3: 53–61.

Easton, H.S. 1978. A comparative study of genetic effects in isogenic diploid and tetraploid plants of Festuca pratensis Huds. 3. Theoretical model. Ann. Amélior. Plantes, 28: 623–636.

Easton, H.S. 1978. A comparative study of genetic effects in isogenic diploid and tetraploid plants of Festuca pratensis Huds. 2. Leaf elongation and seedling growth studies. Ann. Amélior. Plantes, 28: 609–622.

Easton, H.S. 1977. A comparative study of genetic effects in isogenic diploid and tetraploid plants of Festuca pratensis Huds. 1. Introduction, methodology and mean character differences. Ann. Amélior. Plantes, 27: 563–576.

Eisele, C. 1966. What is Chewing's fescue. Saatgutwirtschaft. 18: 342.

Elven, R. 1989. The Omnsbreen Glacier nunataks — a case history of plant immigration. Norw. J. Bot. 27: 1–16.

Evans, G.M., Asay, K.H., and Jenkins, R.G. 1973. Meiotic irregularities in hybrids between diverse genotypes of tall fescue (Festuca arundinacea Schreb.) Crop Sci. 13: 373–379.

Fairey, N.A., and Lefkovitch, L.P. 1993. Agronomic feasibility of producing seed of tall fescue in the Peace River region. Can. J. Pl. Sci. 73: 123–129.

Franklin, J.F., and Dryness, C.T. 1973. Natural vegetation of Oregon and Washington. Oregon State University Press, Corvallis, Oregon, USA.

Fernald, M.L. 1950. Gray's manual of botany. 8th ed. Amer. Book Co., New York, New York, USA.

Flovik, K. 1938. Cytological studies on arctic grasses. Hereditas, 24: 265–376.

Forsling, C.L. and Dayton, W.A. 1931. Artificial reseeding on western mountain rangelands. U.S. Dept. Agric. Circ. 178. 3: 53–61.

Frederiksen, S. 1983. Festuca auriculata in North America. Nord. J. Bot. 3: 632–639.

Frederiksen, S. 1982. Festuca brachyphylla, F. saximontana and related species in North America. Nord. J. Bot. 2: 525–536.

Frederiksen, S. 1981. Festuca vivipara (Poaceae) in the North Atlantic area. Nord. J. Bot. 1: 277–292.

Frederiksen, S. 1979. Festuca minutiflora Rydb., a neglected species. Bot. Not. 131: 409–410.

Frederiksen, S. 1978. Festuca brevissima Jurtz. in Alaska. Bot. Not. 131: 409–410.

Frederiksen, S. 1977. The Festuca brachyphylla group in Greenland. Bot. Not. 130: 269–277.

Gardiner, S.E., Forde, M.B., and Slack, C.R. 1986. Grass cultivar identification by sodium dodeclysulphate polyacrylamide gel electrophoresis. New Zealand J. Agric. Research, 29: 193–206.

Gleason, H.A., and Cronquist, A. 1991. Manual of vascular plants of northeast United States and adjacent Canada. N.Y. Bot. Garden Bronx. New York, USA.

Gould, F.W., and Shaw, R.B. 1983. Grass systematics. 2nd. ed. Texas A & M Univ. Press. College Station, Texas, USA.

Gould, F.W. 1975. The grasses of Texas. Texas A & M Univ. Press. College Station, Texas, USA.

Gower, J.C. 1971. A general coefficient of similarity and some of its properties. Biometrics, 27: 857–871.

Haber, E. 1993. Desktop computer mapping at the National Herbarium of Canada. Taxon, 42: 63–70.

Hackel, E. 1882. Monographia festucarum Europaerum. Theodor Fischer, Berlin, Germany.

Harmer, R. 1984. Vegetative proliferation and vivipary in Scottish grasses. Bot. Soc. Edinb. Trans. 44: 261–268.

Harmer, R., and Lee, J.A. 1978. The growth and nutrient content of Festuca vivipara (L.) Sm. plantlets. New Phytol. 80: 99–106.

Harms, V.L. 1985. A reconsideration of the nomenclature and taxonomy of the Festuca altaica complex in North America. Madroña, 32: 1–10.

Hedberg, O. 1967. Chromosome numbers of vascular plants from arctic and subarctic North America. Ark. Bot. Ser. 2. 6: 309–326.

Hegi, G. 1906. Illustrierte Flora von Mittel-Europa. Verlag von J.F. Lehmann. München, Germany.

Heide, O.H. 1988. Environmental modification of flowering and viviparous proliferation in Festuca vivipara and F. ovina. Oikos, 51: 171–178.

Hickman, J.C. ed. 1993. The Jepson Manual: Higher plants of California. University of California Press, Berkely, Los Angeles, California, USA.

Hitchcock, C.L., Cronquist, A., Ownbey, M., and Thompson, J.W. 1969. Vascular plants of the Pacific Northwest. University of Washington press. Seattle, Washington, USA.

Hitchcock, A.S., and Chase, A. 1951. Manual of grasses of the United States. 2nd ed. U.S. Dep. Agric. Misc. Publ. 200.

Hodgkinson, H.S., and Young, A.E. 1973. Rough Fescue (Festuca scabrella Torr.) in Washington. J. of Range Management, 26: 25–26.

Holmen, K. 1964. Cytotaxonomical studies in the Arctic Alaskan flora. Bot. Not. 117: 109–118.

Holmen, K. 1952. Cytological studies in the flora of Peary Land North Greenland. Medd. om Grønland, 128: 1–40.

Holmgren, P.K., Holmgren, N.J., and Barnett, L.C. 1990. Index Herbariorum. Published for the International Association for Plant Taxonomy, N. Y. Bot. Garden, Bronx, New York, USA

Hooker, W.J. 1840. Festuca. Flora Boreali-Americana Vol. 2. Henry G. Bohn, London, UK. 249–252

Howarth, W.D. 1925. On the occurrence and distribution of Festuca rubra L. sensu amplis, in Britain. J. Linn. London Soc. Bot. 47: 313–331.

Howarth, W.D. 1924. Festuca rubra in Britain. J. Linn. Soc. London Bot. 46: 29–39

Howe, G.M. 1987. Forage cultivar trials — 1987. Agric. Can. Beaverlodge, Alberta, Canada.

Hubbard, C.E. 1968. Grasses: a guide to their structure, identification, uses, and distribution in the British Isles. 2nd ed. Penguin Books. Harmondsworth, Middlesex, UK.

Hultén, E. 1968. Flora of Alaska and neighboring territories. Stanford University Press, Stanford, California, USA.

Hultén, E. 1942. Flora of Alaska and Yukon. 2. Lunds. Univ. Arsskr. N.F., Avd. 2, Bde. 38: 131–412.

Humprhies, C.J. 1980. Lolium. In Flora Europaea, Vol. 5, eds. T.G. Tutin, V.H. Heywood, N.A. Burges, D.M. Moore, D.H. Valentine, S.M. Walters, D.A. Webb. Cambridge University Press, Cambridge, UK.

Hunt, K.L., and Sleper, D.A. 1981. Fertility of hybrids between two geographic races of tall fescue. Crop Sci. 21: 400–404.

Huon, A. 1965. Caractères épidermiques distintifs des ssp. arundinacea (Schreb.) Hack. et pratensis (L.) Hack. du Festuca elatior (L.) Hack. Bull. Soc. Bot. Fr. 112: 37–42.

International Seed Inc. 1996. Golf course professional turf grass specification guide. International Seeds. Halsey, Oregon, USA.

Jarvis, C.E., Stace, C.A., and Wilkinson, M.J. 1987. Typification of Festuca rubra L. Festuca ovina L. and Festuca ovina var. vivipara L. Watsonia, 16: 299–302.

Jauhar, P.P. 1975. Genetic control of dipoid-like meiosis in hexaploid tall fescue. Nature, 254 (5501): 595–597.

Jenkin, T.J. 1959. Fescue species (Festuca L.). In Handbush der Pflanzenzüchtung. eds. H. Kappert and W. Rudorf, Paul Parey, Berlin, Germany. 418–435.

Jenkin, T.J. 1922. Notes on vivipary in Festuca ovina. Bot. Exch. Club. Rep. 6: 418–432.

Johnston, A., Bezeau, L.M., Smith, A.D., and Lutwick, L.E. 1968. Nutritive value and digestibility of fertilized rough fescue. Can. J. Pl. Sci. 48: 351–355.

Johnston, A., and Cosby, H.E. 1966. Rhizomatous form of Festuca scabrella. Can. J. Pl. Sci. 46: 211–212.

Johnston, A., and Bezeau, L.M. 1962. Chemical composition of range forage plants of the Festuca scabrella association. Can. J. Pl. Sci. 42: 105–115.

Johnson, M.C., Pirone, T.P., Siegel, M.R., and Varney, D.R. 1982. Detection of Epichloë typhina in tall fescue by means of enzyme-linked immunosorbent assay. Phytopathology, 72: 647–650.

Johnson, A.W., and Packer, J.G. 1968. Chromosome numbers in the flora of Ogotoruk Creek, N.W. Alaska. Bot. Not. 121: 403–456.

Jordal, L.H. 1952. Some new entities in the flora of the Brooks Range region, Alaska. Rhodora, 54: 35–39.

Jurtzev, B.A., and Zhukova, P.G. 1978. The cytotaxonomical survey of the monocots of the eastern most Chukchi Peninsula. Bot. Zhur. (Lenn.) 63: 1132–1144 (in Russian).

Jurtzev, B.A., and Zhukova, P.G. 1972. Cytotaxonomical characteristics of endemic plants of mountainous north-east Asia. Bot. Zhur. 57: 50–63.

Kerguélen, M. 1975. Les Gramineae (Poaceae) de la flore francaise: essai de mise au point taxonomique et nomenclaturale. Lejeunia ser. 2. 75: 1–343.

Kjellqvist, E. 1961. Studies in Festuca rubra L. Influence of environment. Bot. Not. 114: 403–408.

Kopyto, R., Crane, C.F., and Sleper, D.A. 1989. Effect of temperature on meiosis and fertility in Festuca mairei X F. arundinacea var. glaucescens. Genome, 32: 708–718.

Krajina, V. 1930. Genus Festucam 5. Krajina exposuit. In Schedae ad floram ceschoslovenicam exsiccatum, centuria 2, ed. K. Domin, Acta. Bot. Bohem. 9: 184–220.

Lance, G.N., and Williams, W.T. 1967. A general theory of classificatory sorting strategies. I. Hierarchical systems. Comput. J. 9: 60–64.

Lawrence, J., Hastings, A., Dallwitz, M., and Paine, T. 1993. Beetle larvae of the world. CD-ROM. CSIRO, Canberra, Australia.

Lefkovitch, L.P. 1993. Optimal set covering from biological classification. Agriculture Canada Research Branch, publication 1893/E.

Lewis, E.J. 1977. Studies in Festuca. 4. A phyletic study of Festuca pratensis var. apennina (De Not.) Hack., hybridization with synthetic tetraploid F. pratensis Huds. Genetica, 47: 59–64.

Lewis, E.J. 1963. Hybrids between geographical races of Festuca arundinacea. Schreb. Rep. Welsh Plant Breeding Stn. 1962. 26–27.

Lindenbein, W. 1966. What is Highland Bent and what is Chewings Fescue? Saatgutwirtschaft. 18: 273–274 (in German).

Linder, H.P. 1986. (Poaceae) Diverse notes on southern african pooids. Bothalia, 16: 59–61.

Looman, J. 1982. Grasslands in western North America: Fescue grasslands. In Grassland ecology and classification Symposium Proceedings. eds. A.C. Nicholson, A. McLean, and T.E. Baker. Kamloops, B.C. 209–221.

Looman, J., and Best, K.G. 1979. Budd's flora of the Canadian Prairie Provinces. Agriculture Canada Research Branch, publication 1662.

Looman, J. 1969. The fescue grasslands of western Canada. Vegetatio, 19: 128–145.

Löve, A., and Löve, D. 1975. Cytotaxonomical atlas of the arctic flora. J. Cramer, Vaduz.

Löve, A., and Löve, D. 1956. Cytotaxonomical conspectus of the Icelandic flora. Acta Horti Gotob. 20: 65–291.

Lumaret, R. 1984. The role of polyploidy in the adaptive significance of polymorphism at the GOT 1 locus in the Dactylis glomerata complex. Heredity, 52: 153–169.

Lui, Y., and Dengler, N.G. 1992. Development of open and closed leaf sheaths in Festuca trachyphylla and F. rubra (Poaceae). Can. J. Bot. 70: 1417–1428.

Macoun, J. 1890. Catalogue of Canadian plants. Additions and corrections to parts 1–4. Vol. 2, part 4. Acrogens. Dawson Bros., Montreal, Québec, Canada.

Malik, C.P. 1967. Hybridization of Festuca species. Can. J. Bot. 45: 1025–1029.

Markgraf-Dannenberg, I. 1980. Festuca. In Flora Europaea, Vol. 5, eds. T.G. Tutin, V.H. Heywood, N.A. Burges, D.M. Moore, D.H. Valentine, S.M. Walters, D.A. Webb. Cambridge University Press, Cambridge, UK.

Maude, P.F. 1940. Chromosome numbers in some British plants. New Phytol. 39: 17–32.

McCullagh, P., and Nelder, J.A. 1989. Generalized linear models. 2nd ed. Chapman and Hall, London, UK.

McLean, A. 1970. Plant communities of Similkameen Valley, British Columbia and their relationships to soils. Ecol. Monogra. 40: 403–424.

McLean, A., and Holland, W.C. 1958. Vegetation zones and their relationsips to the soils and climate of the upper Columbia Valley. Can. J. Pl. Sci. 38: 328–345.

McNeill, J., and Dore, W.G. 1976 (1977). Taxonomic and nomenclatural notes on Ontario grasses. Naturaliste Can. (Qué.), 103: 553–567.

Meyers, W.N. 1947. Cytology and genetics of forage grasses. Bot. Rev. 13: 319–421.

Mizianty, M., and Pawlus, M. 1982. Chromosome numbers of some Polish species from the genus Festuca, group Ovina (part 1). Fragmeñta floristica et geobotanica, 28: 363–369.

Mosquin, T., and Hayley, D.E. 1966. Chromosome numbers and taxonomy of some Canadian Arctic plants. Can. J. Bot. 44: 1209–1217.

Muhlenberg, H. 1817. Festuca brevifolia. Descriptio uberior graminum et plantarum calamariarum Americae septentrionalis indigenarum et cicurum. Solomon W. Conrad, Philadelphia, Pennsylvania, USA.

Mulligan, G.A., and Porsild, A.E. 1970. IOPB chromosome number reports 25. Taxon, 19: 102–113.

Murray, J.J., and O'Neill, N.R. 1983. Beltsville 16–1 turf type tall fescue Festuca arundinacea germ-plasm. Crop Sci. 23: 807.

Namzalov, N.N. 1986. The fescue steppes of the southeastern Altai. Izvestiya Sibirskogo otd. Akademii nauk SSSR. Seriya Biol. nauki. 13: 16–24.

Organization for Economic Co-operation and Development. 1993. List of cultivars eligible for certification. OEDC/GD(93)184. Paris.

Page, R.D.M. 1996. TREEVIEW: An application to display phylogenetic trees on personal computers. Computer Applications in the Biosciences 12: 357–358.

Pavlick, L.E. 1985. A new taxonomy of the Festuca rubra complex in northwestern North America, with emphasis on British Columbia. Phytologia, 57: 1–17.

Pavlick, L.E. and J. Looman. 1984. Taxonomy and nomenclature of rough fescues, Festuca altaica, F. campestris (F. scabrella var. major), and F. hallii, in Canada and the adjacent part of the United States. Can. J. Bot. 62: 1739–1749.

Pavlick, L.E. 1984. Studies on the Festuca ovina complex in the Canadian Cordillera. Can. J. Bot. 62: 2448–2462.

Pavlick, L.E. 1983a. Notes on the taxonomy and nomenclature of Festuca occidentalis and F. idahoensis. Can. J. Bot. 61: 337–344.

Pavlick, L.E. 1983b. The taxonomy and distribution of Festuca idahoensis in British Colombia and northwestern Washington. Can. J. Bot. 61: 345–353.

Pavlick, L.E. 1983c. Festuca viridula Vasey (Poaceae): Reestablishment of its original lectotype. Taxon, 32: 117–120.

Pedersen, J.F., Rodriguez-Kabana, R. 1984. Differences among nematode populations in tall fescue Festuca arundinacea pastures in north central and south Alabama, USA. Crop Sci. 24: 819–821.

Pils, G. 1985. Das Festuca vivipara problem in den Alpen. Pl. Syst. Evol. 149: 19–45.

Piper, C.V. 1913. Supplementary notes on American species of Festuca. Contrib. U.S. Natl. Herb. 16: 197.

Piper, C.V. 1906. North American species of Festuca. Contrib. Natl. Herb. 10: 1–48.

Polunin, N. 1940. Botany of the Canadian Eastern Arctic. Part 1. Ottawa, Canada: Natl. Mus. Can., Bull. 92 (Biological series 24).

Porsild, A.E., and Cody, W.J. 1980. Vascular plants of continental Northwest Territories, Canada. Natl. Mus. Nat. Sci., Ottawa, Ontario, Canada.

Porsild, A.E. 1964. Illustrated flora of the Canadian Arctic Archipelago. 2nd ed. Natl. Mus. Can. Bull. 146.

Prentice, H.C., Lönn, M., Lefkovitch, L.P., and Runyeon, H. 1995. Associations between allele frequencies in Festuca ovina and habitat variation in the alvar grasslands on the Baltic island of Öland. J. Ecol. 83: 391–402.

Probatova, N.S. 1976. New and rare species from eastern Siberia and the Far East. Novitates Systematicae Plantarum Vascularum. 13: 37–38.

Prus-Glowacki, W. Sulinowski, S., Nowacki, E. 1971. Immuno-electrophoetic studies of Lolio-Festuca alloploid and its parental species. Biochem. Physiol. Pflanzen (BBP), 162: 417–426.

Ramesmar-Fortner, N.S., Aiken, S.G., and Dengler, N.G. 1995. Phenotypic plasticity in leaves of four species of arctic Festuca (Poaceae). Can. J. Bot. 73: 1810–1823.

Reveal, J.L., and Terell, E.E. 1991. Proposal to reject Festuca elatior L. with comments on the typification of F. pratensis and F. arundinacea (Poaceae). Taxon, 40: 135–137.

Rydberg, P.A. 1917. Flora of the Rocky Mountains and adjacent plains. New York. Published by the author.

Rydberg, P.A. 1909. Studies on the Rocky Mountain Flora. 19. Bull. Torr. Bot. Club, 36: 531–541.

Rydberg, P.A. 1905. Studies on the Rocky Mountain Flora. 15. Bull. Torr. Bot. Club, 32: 597–610.

Rydberg, P.A. 1900. Catalogue of the flora of Montana and the Yellowstone National Park. Festuca vallicola. Mem. N.Y. Bot. Garden 1: 57.

Saint-Yves, A. 1925. Contribution a l'etude des Festuca (subgen. Eu-Festuca) de l'Amerique du Nord et du Mexique. Candollea, 2: 229–316.

Salvesen, P.H. 1986. Festuca vivipara in Southern Norway, infraspecific differentiation and systematic affinities. Acta Univ. Ups., Symb. Bot. Ups. 27: 155–167.

Schmit, R.M., Duell, R.W., and Funk, C.R. 1974. Isolation barriers and self-compatibility in selected fine fescues. Proc. 2nd International Turfgrass Res. Conf. 9–17.

Scoggan, H.J. 1978. The flora of Canada. Part 2. Natl. Mus. Nat. Sci. Publ. Bot. 7: 93–545.

Semikohov, V.F., Kalistratova, O.A., and Are'Feva, L.P. 1981. Variability of the protein fractions of seeds and indices of biochemical evolution. Byull. Mosk. O-Va Ispyt. Prir. Otd. Biol. 86: 100–110.

Shildrick, J.P. 1976. A provisional grouping of cultivars of Festuca rubra L. J. Sports Turf Res. Inst. 52: 9–13.

Siplivinskii, V.N. 1973. Viviparous fescues (Festuca L.) of Siberia outside of the Arctic. Novst. Syst. Vyssh. Rast. 10: 11–18 (in Russian).

Smoliak, S., Adams, B.W., Schuler, B.G., Wroe, R.A., Klump, S.G., and Willms, W.D. 1985. Forage production on selected native prairie sites in southern Alberta. Agric. Can. Tech. Bull. 1985–3E.

Sneath, P.H.A., and Sokal, R.R. 1973. Numerical taxonomy — the principles and practice of numerical classification. W.H. Freeman: San Francisco, California, USA.

Spilsbury, R.H., and Tisdale, E.W. 1944. Soil plant relationships and vertical zonation in the southern interior of British Columbia. Sci. Agric. 24: 395–436.

Stace, C.A., A.-K., Al-Bermani, K.A., and Wilkinson, M.J. 1992. The distinction between the Festuca ovina L. and Festuca rubra L. aggregates in the British Isles. Watsonia, 19: 107–112.

Stace, C.A., and Ainscough, M.M. 1984. Continuing additions to the gene-pool of the Festuca rubra aggregate (Poaceae: Poeae). Pl. Syst. Evol. 147: 227–236.

Staehlin, A., and Tirtapradja, H. 1974. A comparison of reed fescue (Festuca arundinacea) and meadow fescue (Festuca pratensis) from a chemical point of view. Z. Acker-Pflanzenb. 140: 100–116.

Stickney, P.F. 1960. Range of rough fescue (Festuca scabrella Torr.) in Montana. Proc. Montana Acad. Sci. 20: 12–17.

Stringer, W.C., Wolf, D.D., and Blaser, R.E. 1981. Summer regrowth of tall fescue stubble characteristics and micro environment. Agron. J. 73: 96–100.

Swallen, J.R. 1932. Five new grasses from Texas. Amer. J. Bot. 19: 436–442.

Swofford, D.L. 1991. PAUP: phylogenetic analysis using parsimony. Version 3.1. Illinois Natural History Survey. Champaign, Illinois, USA.

Taylor, R.L., and Mulligan, G.A. 1968. Flora of the Queen Charlotte Islands. Part 2. Cytological aspects of the vascular plants. Can. Dep. Agric. Res. Branch Monogr. 4.

Terrell E.E. 1979. Taxonomy, morphology, and phylogeny. In Tall Fescue. eds. R.C. Buckner and L.P. Bush, Amer. Soc. Agronomy, 20. Crop Sci. Soc. Amer. and Soil Sci Soc. Amer. Inc., publishers. Madison. Wisconsin, USA.

Terrell E.E. 1968. Notes on Festuca arundinacea and F. pratensis in the United States. Rhodora, 70: 564–568.

Terrell, E.E. 1967. Meadow Fescue: Festuca elatior L. or F. pratensis Hudson? Brittonia, 19: 129–132.

Tisdale, E.W. 1947. The grasslands of the southern interior of British Columbia. Ecol. 28: 346–382.

Toynbee, K. 1987. Prolific flowering year for Plain Rough Fescue at the Kernen Prairie. Blue Jay, 45: 142–143.

Trehane, P., Brickell, C.D., Baum, B.R., Hetterscheid, W.L.A., Leslie, A.C., McNeill, J., Spongberg, S.A., and Vrugtman, F. (eds.). 1995. International code of nomenclature for cultivated plants — 1995. Quarterjack Publishing, Wimborne, UK.

Tyler, B., Borrill, M., and Chorlton, K. 1978. Studies in Festuca. 10. Observations on germination and seedling cold tolerance in diploid Festuca pratensis and tetrapoid F. pratensis var. apennina in relation to their altitudinal distribution. J. Applied Ecol. 15: 219–226.

Tzvelev, N.N. 1976. Poaceae USSR. Nauka, Leningrad. (in Russian) Translated as Grasses of the Soviet Union Zlaki SSSR. Nauka, Leningrad. Published for Smithsonian Institution Libraries and National Science Foundation. 1983. Oxonian, New Delhi, India.

Tzvelev, N.N. 1972a. On the taxonomy and phylogeny of the genus Festuca L. of the USSR flora. 2. Evolution of the subgenus Festuca. Bot. Zhur. (Lenn.) 57: 161–172 (in Russian).

Tzvelev, N.N. 1972b. Genus Festuca in USSR. Novst. Sist. Vyssh. Rast. 9: 15–46 (in Russian).

Tzvelev, N.N. 1971. On the taxonomy and phylogeny of the genus Festuca L. of the USSR flora. 1. The system of the genus and the main trends of evolution. Bot. Zhur. (Lenn.) 56: 1252–1262 (in Russian).

Vasey, G. 1893a. Descriptions of new or noteworthy grasses from the United States. Contrib. U.S. Natl. Herb. 1: 267–280.

Vasey, G. 1893b. Grasses of the Pacific Slope including Alaska and the adjacent islands. Bull. U.S. Dep. Agric. Div. Bot. 13. Part 2.

Vasey, G. 1876. Notes on Festuca Thurberi. Bot. Gazette. Vol. 2. No. 1.

Villamil, C.B., Duell, R.W., Fairbrothers, D.E., and Sadowski, J. 1982. Isoelectric focusing of esterases for fine fescue identification. Crop Sci. 22: 786–793.

Voss, E.G. 1972. Michigan flora. Part 1. Cranbrook Inst. Sci., Bloomfield, Mich. and Univ. Mich. Herb., Ann Arbor, Michigan, USA.

Watson, L., and Dallwitz, M.J. 1994. The families of flowering plants: interactive identification and information retrieval. CD-ROM. CSIRO, Canberra, Australia.

Watson, L., Gibbs Russell, G.E., and Dallwitz, M.J. 1989. Grass genera of South Africa: interactive identification and information retrieval from an automated data bank. S. Afr. J. Bot. 55: 452–463.

Watson, L., Aiken, S.G., Dallwitz, M.J., Lefkovitch, L.P., and Dubé, M. l986. Canadian grass genera: keys and descriptions in English and French from an automated data bank. Can. J. Bot. 64: 53–70 and 2 microfiche.

Weber, W.A. 1984. New names and combinations, principally in the Rocky Mountain flora. 4. Phytologia, 55: 1–2.

Weber, W.A. 1961. Additions to the flora of Colorado. Univ. Colorado Stud. Ser. Biol. 7: 1–26.

Welsh, S.L., Atwood, N.D., Higgins, L.C., and Goodrich, S. 1987. A flora of Utah. Great Basin Naturalist Memoirs. 9. Brigham Young University. Provo, Utah, USA.

White, J.F. Jr., and Cole, G.T. 1985. Endophyte-host associations in forage grasses 2. Taxonomic observations on the endophyte of Festuca arundinacea. Mycologia, 77: 483–486.

Wilkinson, M.J., and Stace, C.A. 1991. A new taxonomic treatment of the Festuca ovina L. aggregate (Poaceae) in the British Isles. Bot. J. Linn. Soc. 106: 347–397.

Wilkinson, M.J., and Stace, C.A. 1989. The taxonomic relationships and typification of Festuca brevipila Tracey and Festuca lemanii Bastard Poaceae. Watsonia, 17: 289–300.

Woolley, J.B., and Stone, N.D. 1987. Application of artificial intelligence to systematics: SYSTEX — prototype expert system for species identification. Syst. Zool. 36: 246–267.

Woodward, W.U.W., and Frakes, R.V. 1971. Meiotic irregularities in tall fescue genotypes and their F1. hybrids. Oreg. Agric. Exp. Stn. Tech. Bull. 139: 1–56.

Wycherley, P.S. 1953a. Proliferation of spikelets in British grasses. 1. The taxonomy of the viviparous races. Watsonia, 3: 41–56.

Wycherley, P.S. 1953b. The distribution of viviparous grasses in Great Britain. J. Ecol. 41: 275–288.