The genera of Leguminosae-Caesalpinioideae & Swartzieae

L. Watson and M.J. Dallwitz

Moldenhawera Schrader

Moldenhauera Schrad., and cf. Dolichonema Nees

Type species: M. floribunda Schrad.

Habit and leaf form. Trees, or shrubs; unarmed.

Phyllotaxy spiral. The leaves compound; pinnate and bipinnate; with opposite or sub-opposite pinnae; imparipinnate, or paripinnate; with adaxially grooved rachides. The leaflets symmetrical or nearly so; pinnately veined, with a predominant ‘midrib’. Stipules narrow, small, absent or early caducous or very inconspicuous in mature leaves (caducous); membranous. Stipels absent.

Inflorescence and floral morphology. The inflorescences terminal; branched; of racemose units; panicles (comprising subpanicles of racemes). The flowers not distichous. Bracts small and narrow, caducous and absent at anthesis. Bracteoles absent; absent at anthesis.

The flowers hermaphrodite; pentamerous. Hypanthium more or less absent, the androecium hypogynous. The perianth comprising distinct calyx and corolla. Calyx 5; covering the rest of the flower in bud; more or less regular (the segments subequal); members not imbricate (connate at first, later separating valvately). Corolla present; slightly irregular (the spreading, fimbriate petals ovate-oblong, slightly unequal); fimbriate, 5; without greatly reduced members; polypetalous. Petals all clawed; imbricate; imbricate-ascending. Disk absent. The androecium comprising 10 members (with glabrous filaments); not declinate; members all free of one another; members markedly unequal; including staminodia. The staminodia 1; comprising the incurved-ascending abaxial member, which equals the style and is much longer than the fertile stamens. Fertile stamens 9. Anthers sagittate, attached at the base of the connective; dehiscing introrsely. Ovary sessile or subsessile; free. Stigma not dilated (truncate on the filiform style). Ovules numerous.

Fruit, seed and seedling. Fruit a two-valved pod; oblong, compressed, leathery; becoming woody, or not becoming woody. The mature valves without prominent venation. Seeds with a straight or slightly oblique radicle; amyloid-negative.

Transverse section of lamina. Leaves without conspicuous phloem transfer cells in the minor veins. Druses common in the mesophyll. Mesophyll secretory cavities absent. Adaxial hypodermis absent. Leaf girders absent. Laminae dorsiventral. Mesophyll without unaligned fibres or sclereids. Minor veins mainly with abundant accompanying fibres, or lacking accompanying fibrous tissue.

Leaf lamina epidermes. Epidermal crystals not seen either adaxially or abaxially. Simple unbranched hairs common; smooth. Compound or branched eglandular hairs present (T-shaped, in M. cuprea). Capitate glands not seen. Hooked hairs not seen. Cassieae-type leaf pseudo-glands not seen. Expanded and embedded hair-feet absent. Basally bent hairs absent. Adaxial: Adaxial interveinal epidermal cell walls markedly sinuous in high-focus optical section; not conspicuously pitted; thick. Stomata adaxially very rare. Abaxial: Abaxial stomata predominantly paracytic. Abaxial epidermis not papillate. Abaxial interveinal epidermal cell walls straight, or gently undulating, or markedly sinuous in high-focus optical section; conspicuously pitted in optical section; staining normally with safranin; medium-thick.

Pollen ultrastructure. Tectum punctate; smooth punctate, or puncticulate. Length of colpi greater than one half pole to pole distance (without a margocolpus).

Species number and distribution. 13 species. Brazil, Venezuela.

Tribe. Caesalpinieae.

Miscellaneous. Illustrations: • M. floribunda: Fl. Brasiliensis 15 (1870). • M. cuprea: Pohl, Planta Bras. Icones (1887). • M. cuprea leaflet: adaxial epidermis. • M. cuprea leaflet: abaxial epidermis.

We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, distributions of character states within any set of taxa, geographical distribution, and classification. See also Guidelines for using data taken from Web publications.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1993 onwards. The genera of Leguminosae-Caesalpinioideae and Swartzieae: descriptions, illustrations, identification, and information retrieval. In English and French. Version: 22nd March 2017.’.