The Genera of Leguminosae-Caesalpinioideae and Swartzieae

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L. Watson and M. J. Dallwitz

Daniella J.J. Benn

Habit and leaf form. Trees, or shrubs; without tendrils; unarmed. Phyllotaxy spiral. Leaves compound; pinnate; paripinnate. Leaflets many per leaf; symmetrical or nearly so at their bases; opposite or sub-opposite, or alternate (rare); without a continuous marginal nerve; petiolulate; with petiolules not noticeably twisted. Stipules absent or early caducous or very inconspicuous; neither leafy nor spinescent; not connate (intrapetiolar, enfolding bud). Stipels absent.

Inflorescence and floral morphology. Flowers hermaphrodite; not pentamerous throughout; departing from pentamery in the calyx; coloured; in panicles (of racemes); not distichous. Inflorescences terminal; of racemose units. Bracts absent at anthesis. Bracteoles present; relatively large and enclosing the flower buds; absent at anthesis, or persistent beyond anthesis; not valvate. Hypanthium present. Length of floral tube relative to total hypanthium plus calyx length, about 0.25. Calyx polysepalous; covering the rest of the flower in bud, or not covering the rest of the flower in bud; more or less regular; 4 partite (deciduous); imbricate; not Swartzieae type. Corolla present; very zygomorphic; polypetalous; including greatly reduced petals. Petals 5; imbricate; imbricate-ascending. Clawed petals absent. Disk absent. Androecium of ten parts; with united members, or members all free of one another; members all more or less equal in length; without staminodia. Fertile stamens 10. Anthers attached well above base of connective. Dehiscence longitudinal. Ovary stipitate; excentric with adnate stipe. Ovules numerous.

Fruit, seed and seedling. Fruit a two-valved pod; not becoming woody; curved; without prominent raised veins; not winged; valves twisting and enrolling during dehiscence. Seeds non-endospermic; arillate (with cupular aril and long funicle); with a straight or slightly oblique radicle; amyloid-positive. Cotyledons flat; of Type 4; with a vascular system in one plane; epigeal.

Transverse section of lamina. Leaves with conspicuous phloem transfer cells in the minor veins. Druses common in the mesophyll, or absent from the mesophyll. Mesophyll secretory cavities (gland-dots) common; epithelium-lined. Adaxial hypodermis absent. Leaf girders absent. Laminae dorsiventral. Mesophyll without unaligned fibres or sclereids. Minor veins mainly with abundant accompanying fibres.

Leaf lamina epidermes. Epidermal crystals present; prisms, or druses. Simple unbranched hairs common, or not seen. No compound or branched eglandular hairs seen. Capitate glands not seen. Hooked hairs not seen. Cassieae-type leaf pseudo-glands not seen. Expanded and embedded hair-feet absent. Adaxial interveinal epidermal cell walls straight in optical section; conspicuously pitted, or not conspicuously pitted; thick. Stomata adaxially very rare. Abaxial stomata predominantly paracytic. Abaxial epidermis not papillate. Abaxial interveinal epidermal cell walls straight, or gently undulating; conspicuously pitted in optical section; staining normally with safranin; thick.

Wood anatomy. Wood without septate fibres; storied; with normal intercellular canals; without traumatic canals. Intervascular pits medium to large.

Pollen ultrastructure. Tectum punctate; rugulose punctate. Length of colpi greater than one half pole to pole distance.

Cytology, geography, etc. Basic chromosome number, x = 11. 2n = 22. 9 species. Tropical West Africa. Not widely cultivated.

Tribe. Detarieae.


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Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1993 onwards. The genera of Leguminosae-Caesalpinioideae and Swartzieae: descriptions, identification, and information retrieval. In English and French; French translation by E. Chenin. Version: 19th October 2005. http://delta-intkey.com’.

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