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The moss families (Bryophyta, Musci) of Britain and Ireland

L. Watson and M.J. Dallwitz

Sematophyllaceae

Signal-mosses.

~Hypnaceae

Gametophyte. Pleurocarpous; slender mosses, forming patches (these tight, glossy and yellowish-tinged), or growing through other Bryophytes. Primary stems procumbent. Shoots (sub-) complanate, or not complanate. Paraphyllia absent. Pseudoparaphyllia absent. Stems yellowish, or green. The leaves of main stems and branches similar in form; bilaterally symmetrical (to only slightly asymmetrical); ovate, or lanceolate; spiral; (sub-) secund, or not secund. Leaf bases not decurrent. The leaves nerveless, or double-nerved, or single-nerved (sometimes, in S. demissa). The leaf nerves when present, not extending beyond the middle of the leaf ("nerve absent or short and double" according to British authors, although their own descriptions represent leaves of S. demissa as "nerveless or with faint traces of a single or double nerve"); without stereids. Leaf blade apices pointed; apically acute to acuminate. Leaf blade margins flat, or revolute or recurved (proximally); unistratose; distally denticulate, or entire. Leaf blades not conspicuously bordered. The basal leaf cells longitudinally much elongated; rhomboidal to linear, or vermicular; smooth. The walls of basal leaf cells straight. The angular cells clearly differentiated (these large, forming small auricles). The mid-leaf cells longitudinally much elongated; rhomboidal to linear, or vermicular; smooth. The walls of the mid-leaf cells straight.

Plants monoecious, or dioecious; when monoecious, autoecious.

Sporophyte. Capsules exserted; inclined to horizontal; symmetrical; straight; ellipsoid; without an externally conspicuous apophysis; smooth; without an annulus. Calyptra symmetrical; splitting down one side. Capsules with a peristome. The peristome double. The peristome teeth 16; not grouped; not deeply cleft (?); not perforated; thin, membranous, and transversely barred; exteriorly with a fine longitudinal dividing line between the transverse bars. The inner peristome well developed, or reduced; with a basal membranous ring; with elongated “processes”. The processes of the inner peristome alternating with the teeth of the outer peristome. The inner peristome ciliate. The operculum rostrate. Setae long; flexuose; smooth, or rough.

Ecology. In wet places to mesophytic. On lightly to moderately shaded rocks in humid habitats, at low altitude (S. demissum), or to 500 m (S. micans).

Representation in Britain and Ireland. 3 species. Sematophyllum. Northern Scotland, southern Scotland, northern England, Wales, and Ireland.

Classification. Class Bryopsida; Subclass Bryideae; Order Hypnales.

Illustrations. • Sematophyllum demissum and S. micans: Dixon. SEMATOPHYLLACEAE: H, Sematophyllum demissum (Wils.) Mitt.; I, Sematophyllum micans (Mitt.) Braithw.

From Dixon and Jameson (1924), the unscaled screen display approximately doubling the magnifications given with their individual figures. 1, leaf, or stem leaf where these differ; 2, branch-leaf; 3, perichaetial leaf; 4, perigonial leaf; 5, capsule; 6, peristome; 7, calyptra; 8, spores; 9, inflorescence; 10, gemmae; 11, paraphyllia; 12, stoma of capsule; *, plant, or part of one. Qualifications: a, apex; b, base; c, cells at one third from the apex; cv, ventral aspect of cells; cd, dorsal aspect of cells; bc, basal cells; x, section. • Sematophyllum demissum, with Campyliaceae and Hypnaceae: Berkeley. SEMATOPHYLLACEAE: 6, Sematophyllum demissum (Wils.) Mitt., habit; a, abaxial (above) and adaxial leaves; b, capsule. CAMPYLIACEAE: 1, Drepanocladus uncinatus (Hedw.) Warnst., habit; b, capsule. (cf. Berkeley’s Plate 10,2.) 5, Scorpidium scorpioides (Hedw.) Limpr., habit; a, leaf; b, capsule; c, male inflorescence. HYPNACEAE: 2, Ptilium cristacastrensis (Hedw.) De Not, habit; a, leaf; b, capsule. 3, Ctenidium molluscum (Hedw.) Mitt., unusual habit form; a,leaves; b, capsule. 4,Hypnum cupressiforme Hedw., habit; a, leaf; b, capsule.

From Berkeley (1863, Plate 11). The approximate 1 cm scale applies to the whole plant (habit) illustrations only.


We advise against extracting comparative information from the descriptions. This is much more easily achieved using the DELTA data files or the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa. See also Guidelines for using data taken from Web publications.


Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 2005 onwards. The moss families (Bryophyta, Musci) of Britain and Ireland. Version: 5th August 2019. delta-intkey.com’.

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