British insects: the families of Coleoptera

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L. Watson and M. J. Dallwitz

Heteroceridae

Mud-beetles.

Adults. Beetles terrestrial, or terrestrial to sub-aquatic (some are found in wet - including brackish - habitats); walking in water or free-swimming by conventional ambulatory motion of the legs, not diving strongly; perhaps replenishing air for respiration beneath the elytra indirectly from that acquired and held in a ventral plastron (?); incorporating bubbles of oxygen directly into the plastron (?). Beetles 2.5–5 mm long; body length/maximum body width 1.7–3.05; elytral length/pronotal length 2.2–3.53; base of prothorax not or scarcely narrower than the combined elytral bases; greatest prothoracic width not narrower or only slightly narrower than the greatest elytral width. Beetles elongate; without ventral body cavities for reception of the legs; not necked; conspicuously waisted. Upper surfaces of body non-glabrous; not bristly; with neither scales nor scale-like setae. Beetles prognathous. Inclination of the head slight.

Eyes not strongly protuberant; without bristles; finely facetted. Mandibles with a well developed mola. The mandibular apices simple, or bidentate or bilobed. The incisor edges of the mandibles with a single tooth, or with two or more teeth. The maxillae with distinct galea and lacinia apically to the palp. The apical segment of the maxillary palps cylindrical to fusiform. The apical segment of the labial palps not expanded apically. Antennae very short; strongly asymmetric; (9–)11 segmented. Antennal scape not swollen. Antennae clubbed. Antennal clubs 7 segmented (serrate); preceded by a cupule, or without a cupule. Antennal insertions visible from above; not in fossae.

Prothorax shorter than wide; at its widest not markedly narrower than the adjoining part of the abdomen; without notopleural sutures. Pronotal length/maximum pronotal width 0.52–0.82. Metaventrite with a transverse groove. The fore-leg coxal cavities broadly open; narrowly separated; strongly transverse; without lateral extensions; internally open. Hind coxae posteriorly shaped to receive the retracted femur; with a steep transverse declivity against which the femur retracts. Tarsal segmentation formula 4, 4, 4. None of the tarsi with conspicuously bilobed segments. None of the tarsi with ‘hidden’ segments (but the terminal segment about half as long as the basal three together). The front tarsi with as many segments as the mid-tarsi; 4-segmented. Mid-leg tarsi 4-segmented; tetramerous. The claws of the mid-leg tarsi not appendaged. The claws of the mid-leg tarsi simple. Hind tarsi with at least as many segments as the mid-tarsi; 4-segmented.

Elytral length/maximum width across the elytra 1.18–2.3. Elytra exposing no more than part of the terminal tergite; not truncate; hard. Scutellary striole absent. Elytra non-glabrous (densely hairy). Wings well developed, or absent or much reduced. Abdominal tergites 10. Exposed abdominal sternites 5; comprising both fused and movable components; immovably joined 3. Abdominal segment 8 with apparently functional spiracles.

Adults not predacious; phytophagous; in living vegetation, or in decaying plant material (found in stiff mud at the sides of ponds and streams).

Larvae. The larvae campodeiform; elongate and more or less parallel-sided; vestiture restricted to fine hairs or setae; dorsally heavily pigmented or sclerotized; only very lightly pigmented. The antennae 3 segmented; less than 0.15 x the with of the head. Stemmata fewer than 6 (five). Frontoclypeal suture between frons and clypeus distinct. The labrum and head capsule separated by a complete suture. Apices of the mandibles bilobed or bidentate. The maxillary palps 3 segmented. The labium with a short ligula between the palps. The labial palps 3 segmented. The mesothoracic legs 5 segmented (including the pretarsus); with 1 movable claw. Visible abdominal segments 10. Tergum 9 of the abdomen entirely dorsal.

The larvae seemingly non-aquatic. The abdomen exhibiting functional abdominal spiracles; having functional spiracles on anterior segments. The larvae without gills. Abdominal tergum 8 without amature. The larvae not predacious; phytophagous; on living vegetation, or in decaying plant material (in mud).

The larvae campodeiform, elongate, with rather distended thoracic segments, well developed legs, and 10 visible abdominal segments.

Worldwide and British representation. About 250 species worldwide; genera 5–20 (depending on taxonomic opinion). 8 species in Britain; genera in Britain 1; Heterocerus.

Classification. Suborder Polyphaga; Infraorder Elateriformia; Superfamily Dryopoidea. E.g., Heterocerus obsoletus (Largest Heterocerus).

Miscellaneous. • Heterocerus obsoletus Curtis (Largest Heterocerus: B. Ent. 224). • Heterocerus obsoletus (details, B. Ent. 224). • Heterocerus obsoletus (B. Ent. 224, legend+text). • Heterocerus obsoletus (B. Ent. 224, text cont.). • Heterocerus fenestratus, H. flexuosus, H. fusculus and H. maritimus: (with Elmidae and Dryopidae): Fowler 3, 98 (1889). • Fowler 3, 98 (1889): original legend.. • H. flexuosus, H. marginatus, H. obsoletus (from joy, 1932).


To view the illustrations with detailed captions, go to the interactive key. This also offers full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 2003 onwards. British insects: water beetles. Version: 18th September 2012. http://delta-intkey.com’.

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