Insects of Britain and Ireland: the families of Coleoptera

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L. Watson and M. J. Dallwitz

Nemonychidae

Cimberididae, Rhinomaceridae.

Pine Weevils.

General appearance. 3–5 mm long. Body length/maximum body width 2.5–3.4. Elytral length/pronotal length 2.25–3.15. Base of prothorax distinctly narrower than the combined elytral bases. Greatest prothoracic width distinctly narrower than greatest elytral width. Body not noticeably widest either behind the thorax, or at the rear (the elytra parallel-sided). Beetles elongate; not necked; conspicuously waisted; neither particularly long- nor short-legged; pitchy black, covered with light yellow or orange hairs; exhibiting bright ‘warning colours’. Upper surfaces of body non-glabrous; not bristly; with neither scales nor scale-like setae.

Detailed morphology. Beetles equipped with a rostrum (this long). Inclination of the head slight. Eyes strongly protuberant; bristly, or without bristles. The labrum at least partly visible in antero-dorsal view (distinguishable as a separate segment from the clypeus); labrum mostly moderately to heavily sclerotized. Mandibles with a reduced mola; with reduced prosthecae, or without prosthecae. The mandibular apices simple. The incisor edges of the mandibles simple, or with a single tooth. The maxillae with distinct galea and lacinia apically to the palp. The maxillary palps normal, flexible; apical segment of the maxillary palps cylindrical to fusiform. The apical segment of the labial palps not expanded apically. Antennae about half the insect's head to tail length to long, but not exceeding the insect’s head to tail length; not elbowed; 11 segmented; not hairy. Antennal scape not swollen. Antennae gradually expanding towards the apex. Antennal insertions hidden from above.

Cervical sclerites present. Prothorax about as long as wide. Pronotal length/maximum pronotal width 0.8–1.2. The pronotum without lateral keels. Prothorax at its widest markedly narrower than the adjoining part of the abdomen. Prothorax without notopleural sutures. Scutellum conspicuous; not elevated; anteriorly simple; posteriorly broadly rounded or obtusely angulate, or truncate. The prosternal process present; interrupted, or entire; when not interrupted, incomplete; when not interrupted, falling short of the mesoventrite to slightly overlapping the mesoventrite. Metaventrite without a transverse groove. The fore-leg coxae countersunk in ‘procoxal cavities’. The fore-leg coxal cavities closed behind externally; narrowly closed; medianly confluent; circular to longer than wide; without lateral extensions; broadly closed internally. The mid-leg coxae countersunk in ‘mesocoxal cavities’; separated by less than the shortest diameter of the cavity. The mid-leg coxal cavities narrowly separated; not or scarcely oblique; closed laterally. Hind-leg coxae contiguous or narrowly separated; extending laterally to meet the elytra, or not markedly extended laterally; not shaped posteriorly to receive the femur. Tarsal segmentation formula 4, 4, 4. The tarsi exhibiting bilobed segments; with a tiny penultimate segment hidden by distal lobing of the fourth and fused to the fifth. Front tarsi with as many segments as the mid-tarsi; 4-segmented, or 5-segmented (depending on interpretation). Mid-leg tarsi 4-segmented, or 5-segmented; pseudotetramerous; the penultimate segment distinctly shorter than the antepenultimate one. The claws of the mid-leg tarsi not appendaged. The claws of the mid-leg tarsi simple, or one-toothed or bifid. Hind tarsi with as many segments as the mid-tarsi; 4-segmented, or 5-segmented.

Elytral length/maximum width across the elytra 1.5–2.05. Elytra exposing no more than part of the terminal tergite. The pygidium entirely concealed by the ends of the elytra even when viewed from behind. Elytra not truncate; non-glabrous (with yellow or orange hairs); without striae. Elytra with six or more longitudinal lines of punctures, or apunctate, irregularly punctate, or each with fewer than 6 longitudinal lines of punctures or impressed striae; the lines per elytron when more than five, 10, or 11. Scutellary striole present. Wings well developed, or absent or much reduced. Wings with an anal lobe. Wings without a medial fleck. Abdominal sternites 5; all articulated and moveable. Abdominal segment 8 apparently without functional spiracles. The male external genitalia cucujiform.

Adult habitat, ecology. On living vegetation (on conifers); phytophagous (on pollen of conifers).

Larvae. Mature larvae minute (less than 3 mm long), or small to medium-sized. The larvae elongate and more or less parallel-sided; C-shaped in lateral view, or not C-shaped. Body circular in cross-section (mostly), or somewhat flattened. Vestiture restricted to fine hairs or setae. The larvae dorsally only very lightly pigmented or sclerotized; ventrally only very lightly pigmented. The antennae 1 segmented. Stemmata present; on either side of the larval head 1. The frontoclypeal suture between frons and clypeus indistinct or absent. The labrum and head capsule partly fused, with an incomplete suture. Apices of the mandibles bilobed or bidentate. The maxillary palps 3 segmented. The labium without ligula between the palps. Labial palps present and segmented; 2 segmented. Mesothoracic legs present and segmented; 1 segmented; with 1 moveable claw. Visible abdominal segments 10. Tergum 9 of the abdomen entirely dorsal. The abdomen having functional spiracles on anterior segments (these annular-biforous); without spiracular tubes. The abdominal apex without a respiratory chamber. Abdominal tergum 8 without amature. The last abdominal segment without cerci.

Larvae in living vegetation; phytophagous (associated with pollen of softwoods).

Classification. Suborder Polyphaga; Superfamily Curculionoidea.

Representation in Britain and Ireland, and worldwide. Genera about 25 (?). 1 species in Britain (C. attelaboides); genera in Britain 1; Rhinomacer, = Cimberis.

General comments. Adults with the labrum separate from the clypeus.

Illustrations. • Cimberis (Rhinomacer) attelaboides (from Joy). Cimberis attelaboides. • Cimberis attelaboides (with Anthribidae and Attelabidae): Fowler 5, 152 (1891). • Fowler 5, 152 (1891): original legend..


To view the illustrations with detailed captions, go to the interactive key. This also offers full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 2003 onwards. Insects of Britain and Ireland: the families of Coleoptera. Version: 16th May 2016. delta-intkey.com’.

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