Insects of Britain and Ireland: the families of Coleoptera

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L. Watson and M. J. Dallwitz



General appearance. 1–1.3 mm long. Body length/maximum body width 2.4–2.6. Elytral length/pronotal length 2.2–2.5. Base of prothorax distinctly narrower than the combined elytral bases. Greatest prothoracic width distinctly narrower than greatest elytral width. Beetles elongate; dorsally flattened to dorsally somewhat convex; not necked; conspicuously waisted; dull black. Upper surfaces of body non-glabrous; not bristly; with neither scales nor scale-like setae.

Detailed morphology. Beetles without a rostrum; prognathous. Inclination of the head slight. Eyes strongly protuberant (as illustrated in H. latridioides); without bristles; coarsely facetted. Ocelli absent. The labrum at least partly visible in antero-dorsal view; labrum mostly moderately to heavily sclerotized. Mandibles with a well developed mola; with well developed prosthecae. The mandibular apices bidentate or bilobed. The incisor edges of the mandibles with a single tooth. The maxillae with distinct galea and lacinia apically to the palp. Antennae short; 11 segmented; rather weakly clubbed. Antennal clubs 3 segmented. Antennal insertions visible from above.

Cervical sclerites absent. Prothorax about as long as wide. Pronotal length/maximum pronotal width 0.95–1.1. The pronotum without lateral keels. Prothorax at its widest markedly narrower than the adjoining part of the abdomen (rather straight-sided). Prothorax without notopleural sutures. Scutellum conspicuous; elevated above the mesoscutum in lateral view; anteriorly simple; posteriorly broadly rounded or obtusely angulate, or truncate. The prosternal process complete; slightly overlapping the mesoventrite. Metaventrite without a transverse groove. The fore-leg coxal cavities quite widely separated. The mid-leg coxae countersunk in ‘mesocoxal cavities’; separated by less than the shortest diameter of the cavity. The mid-leg coxal cavities moderately to widely separated; not or scarcely oblique; closed laterally. Hind-leg coxae not much enlarged; not markedly extended laterally. Tarsal segmentation formula 4, 4, 4. The tarsi without bilobed segments; without ‘hidden’ segments. Front tarsi with as many segments as the mid-tarsi; 5-segmented. Mid-leg tarsi 5-segmented; pentamerous; the penultimate segment not distinctly shorter than the antepenultimate one. The claws of the mid-leg tarsi not appendaged. The claws of the mid-leg tarsi simple. Hind tarsi with as many segments as the mid-tarsi; 5-segmented.

Elytral length/maximum width across the elytra 1.6–1.8. Elytra covering most to all of the abdomen; exposing no more than part of the terminal tergite (most of it). The pygidium at least partly exposed beyond the long elytra. Elytra truncate; dull; non-glabrous. Elytra apunctate, irregularly punctate, or each with fewer than 6 longitudinal lines of punctures or impressed striae. Scutellary striole absent. Elytra absent or incomplete. Wings well developed; fringed with long hairs on the hind margin. Wings without an anal lobe. Abdominal sternites 5; all articulated and moveable. Abdominal segment 8 apparently without functional spiracles. The male external genitalia cucujiform.

Adult habitat, ecology. Land-dwellers; not predacious (?); associated with dung.

Larvae. Mature larvae minute (less than 3 mm long). Body circular in cross-section, or somewhat flattened (?).

Larvae in dung.

Classification. Suborder Polyphaga; Superfamily Cucujoidea.

Representation in Britain and Ireland, and worldwide. 1 species worldwide (to several?); genera 1 (Hypocoprus). 1 species in Britain; genera in Britain 1; Hypocoprus. E.g., H. latridioides.

General comments. Seemingly restricted to the northern hemisphere, and poorly known. Unwin’s citation of tarsi as 4,4,4 is contradicted by Lawrence et al.

Illustrations. • Hypocoprus latridioides (from Joy). Hypocoprus latridioides. • Hypocoprus latridioides: Fowler 3, 93 (1889).

To view the illustrations with detailed captions, go to the interactive key. This also offers full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 2003 onwards. Insects of Britain and Ireland: the families of Coleoptera. Version: 16th May 2016.’.