British ferns (Filicopsida)

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L. Watson and M. J. Dallwitz

Dryopteris remota (A. Braun ex Döll) Druce

“Scaly Buckler Fern”.

D. woynarii Rothm.

Sporophyte. Plants rhizomatous. The rhizomes stout; bearing scales. Plants with no clear distinction into fertile and sterile leaves.

Leaves aggregated terminally; to 20–75 cm long; circinnate; laminate; compound; complexly divided; bipinnate with conspicuously divided pinnules (the pinnules evenly lobed at least to halfway, or almost to the midrib - this supposedly distinguishing D. remota from bipinnate D. affinis). Pinnae 15–25 on each side of the leaf (?). The petioles shorter than the blades (with dense golden brown scales, these often with darker centres); about 0.3–0.5 x the length of the blade (about 1/3 to half as long); not jointed; vascularised by several discrete strands (derived via several leaf traces). Petiolar scales golden brown and often dark centred, dense. Leaf blades in outline narrowly ovate; not thin and translucent. The longest pinnae near the base of the blade, or about a third of the distance from the base of the blade, or around the middle of the blade (? - the lowest pinnae about 4/5 the length of the longest). The pinnae decreasing markedly in length towards the base of the blade, the basal ones relatively short to not decreasing markedly in length basipetaly, the basal pinnae not or scarcely shorter than the longest pair. The venation of the lamina open.

Homosporous; leptosporangiate. The sporangia tiny and delicate, stalked, all alike, borne in aggregates; superficial; protected; aggregated in sori. The sori sub-orbicular; remaining discrete at maturity; with a true indusium. The indusia reniform and attached at the indentation. Paraphyses absent. The sporangia not developing sequentially. The sporangial wall of only one cell layer. The mature spores monolete; without a perispore.

Prothallus. Prothalli free-living.

Distribution and habitat. Formerly in N. Kerry, S.E. Galway and possibly Dunbarton, seemingly now extinct save in cultivation.

Vice-county records. Britain: Herefordshire, Stirlingshire, Dunbartonshire. Ireland: South-east Galway.

Classification. Family Polypodiaceae (C.T.W.); Dryopteridaceae (Swale and Hassler); Dryopteridaceae (Stace). Order Dryopteridales (Swale and Hassler).

Comments. An apogamous hybrid derivative of D. affinis x D. expansa.

Illustrations. • D. remota: as Lastrea remota, Eng. Bot. 1852 (1886).


To view the illustrations with detailed captions, go to the interactive key. This also offers full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting or lacking specified attributes, and distributions of character states within any set of taxa.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 2004 onwards. British ferns (Filicopsida). Version: 4th January 2012. http://delta-intkey.com’.

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