The Families of Flowering Plants | |
~ Fumariaceae, Papaveraceae sensu lato
Habit and leaf form. Stemless herbs (with Blechnum-like leaves); non-laticiferous and without coloured juice. Perennial; with a basal aggregation of leaves (the leaves all radical); rhizomatous. Leaves alternate (surrounded at the base by several large, orbicular, membranous cataphylls); simple, or compound; if considered compound, very evenly pinnate. Lamina if considered entire, very deeply dissected; regularly, evenly pinnatifid. Leaves exstipulate.
General anatomy. Plants without laticifers.
Leaf anatomy. Stomata present; mainly confined to one surface (the abaxial); anomocytic.
The mesophyll without sclerenchymatous idioblasts.
Stem anatomy. Secondary thickening absent.
Reproductive type, pollination. Fertile flowers hermaphrodite. Plants hermaphrodite.
Inflorescence, floral, fruit and seed morphology. Flowers aggregated in inflorescences; in fascicles. Inflorescences scapiflorous (the scape erect, simple or very slightly branched); (scapes) bearing more or less distant, two-flowered fascicles of flowers on slender pedicels. Flowers regular to somewhat irregular (almost regular); cyclic. Free hypanthium absent.
Perianth with distinct calyx and corolla; 6; 3 whorled (2+2+2); isomerous. Calyx 2; 1 whorled; polysepalous; neither appendaged nor spurred (and the sepals not lobed); not persistent (caducous); open in bud (not enclosing the flower bud). Corolla 4; 2 whorled (2+2); polypetalous; unequal but not bilabiate, or regular (the petals subequal); white; deciduous (caducous). Petals entire.
Androecium ostensibly 4. Androecial members unbranched; free of the perianth; free of one another; 1 whorled. Androecium exclusively of fertile stamens. Stamens 4; oppositisepalous; alternating with the corolla members (diagonal); very shortly filantherous (without basal nectaries). Anthers oblong; all bilocular; tetrasporangiate (?). Pollen shed as single grains. Pollen grains aperturate; (2–)3(–4) aperturate; colpate.
Gynoecium 2 carpelled. Carpels isomerous with the perianth. The pistil 1 celled. Gynoecium syncarpous; synstylovarious; superior. Ovary 1 locular (flattened). Gynoecium stylate. Styles 1; apical. Stigmas 2; commissural. Placentation basal. Ovules in the single cavity 2–4; non-arillate (at least, the seeds neither strophiolate nor carunculate); anatropous, or anatropous to campylotropous; bitegmic; crassinucellate. Outer integument not contributing to the micropyle. Endosperm formation nuclear.
Fruit non-fleshy; dehiscent; capsular-indehiscent. Capsules (two) valvular (siliculiform, dehiscence acropetal). Fruit many-seeded.
Physiology, biochemistry. Alkaloids present.
Geography, cytology. Holarctic. Temperate. Japan. N = 9.
Taxonomy. Subclass Dicotyledonae; Crassinucelli. Dahlgrens Superorder Ranunculiflorae; Papaverales (?). Cronquists Subclass Magnoliidae; Papaverales. APG 3 core angiosperms; peripheral eudicot; Superorder Ranunculanae; Order Ranunculales (as a synonym of Papaveraceae).
Species 1. Genera 1; only genus, Pteridophyllum.
Illustrations. • Pteridophyllum (Chittenden).
This description is offered for casual browsing only. We strongly advise against extracting comparative information from it. This is much more easily achieved using the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting specified attributes, summaries of attributes within groups of taxa, geographical distribution, genera included in each family, classifications (Dahlgren; Dahlgren, Clifford, and Yeo; Cronquist; APG), and notes on the APG classification.
Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1992 onwards. The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Version: 25th November 2009. http://delta-intkey.com’.