The Families of Flowering Plants

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L. Watson and M. J. Dallwitz

Implicit Attributes

Unless indicated otherwise, the following attributes are implicit throughout the descriptions, except where the characters concerned are inapplicable.

Habit and leaf form. Non-laticiferous and without coloured juice. ‘Normal’ plants. Leaves well developed. Plants with roots; non-succulent; autotrophic. Young stems not breaking easily at the nodes. Rhizome and root tissues not red-pigmented. Self supporting. Stem growth not conspicuously sympodial. Non-marine. Leaves not connate. Leaf sheaths with free margins. Leaves not gland-dotted; without marked odour; with ‘normal’ orientation; not peltate. Lamina neither inverted nor twisted through 90 degrees; not conspicuously asymmetric. Leaves eligulate. Stipules not ochreate; without colleters. Axillary scales absent. Vernation not circinnate. Leaves becoming compound from primordial lobes. Domatia never explicitly mentioned for the family.

General anatomy. Plants without laticifers. Plants without ‘crystal sand’. Plants without silica bodies.

Leaf anatomy. Leaves without ‘pearl glands’. Epidermis without differentiation into ‘long’ and ‘short’ cells; without silica bodies. Guard-cells not ‘grass type’. Urticating hairs absent.

Lamina without secretory cavities. Cystoliths absent. The mesophyll without etherial oil cells; not containing mucilage cells; without sclerenchymatous idioblasts.

Stem anatomy. The cortex without cristarque cells. Cortical bundles absent. Medullary bundles absent. Internal phloem absent. The secondary phloem not stratified. Xylem without tracheids; without fibre tracheids; without libriform fibres; with vessels. Tile cells absent. Pith without diaphragms.

Root anatomy. Roots without velamen.

Inflorescence, floral, fruit and seed morphology. The fruiting inflorescences not conelike. Flowers not in ‘spikelets’. Inflorescences not scapiflorous; without involucral bracts; not pseudanthial; espatheate. Bracts not calyptrate. Flowers not calyptrate; not resupinate; not exhibiting the androecium internal to the gynoecium; neither papilionaceous nor pseudo-papilionaceous; cyclic. Floral receptacle with neither androphore nor gynophore.

Perianth without spots. Calyx not represented by bristles; neither appendaged nor spurred; not calyptrate. Epicalyx absent. Corolla alternating with the calyx; not appendiculate. Corolla tube not noticeably adaxially split. Corolla not calyptrate; not spurred; not fleshy.

Androecial members unbranched; free of the gynoecium. Androecium exclusively of fertile stamens. Staminodes non-petaloid. Stamens not didynamous, not tetradynamous; filantherous. Filaments not appendiculate. Anthers separate from one another; not becoming inverted during development; bilocular; unappendaged. Pollen shed as single grains. Pollen grains not dispersed as long filaments. The ulcus without an operculum; without an annulus.

Gynoecium non-petaloid. Carpel fully closed. Styles free. Locules without ‘false septa’; not horizontally divided. Styles without an indusium. Stigmas dorsal to the carpels.

Fruit not an aggregate. The fruiting carpels not coalescing. Fruit without fleshy investment. Gynoecia of adjoining flowers not forming a multiple fruit. The multiple fruits not coalescing. Endosperm not ruminate. Perisperm absent. Seeds not conspicuously hairy; with a testa; wingless; without amyloid. Embryo well differentiated. Testa without phytomelan. Micropyle not zigzag.

Seedling. Coleoptile absent. Primary root persistent.

Physiology, biochemistry. Nitrogen-fixing root nodules absent. Mustard-oils absent. Polyacetylenes not found. Berberine absent. Andromedotoxin not recorded. Betalains absent. Inulin not found.

Peculiar feature. Gynoecium and fruit not as in Resedaceae. Lamina tip not abaxially pouched. Epicalyx not double and complex as in Triplostegiaceae.

Geography, cytology. Chromosomes with normal centromeres.


This description is offered for casual browsing only. We strongly advise against extracting comparative information from it. This is much more easily achieved using the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting specified attributes, summaries of attributes within groups of taxa, geographical distribution, genera included in each family, classifications (Dahlgren; Dahlgren, Clifford, and Yeo; Cronquist; APG), and notes on the APG classification.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1992 onwards. The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Version: 25th November 2009. http://delta-intkey.com’.

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