The Families of Flowering Plants

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L. Watson and M. J. Dallwitz

Gyrocarpaceae Dum.

~ Hernandiaceae

Habit and leaf form. Small trees, or shrubs, or lianas. Self supporting, or climbing. Mesophytic. Leaves alternate; petiolate; non-sheathing; simple. Lamina dissected, or entire; when dissected, palmatifid; palmately veined; cross-venulate. Leaves exstipulate; without a persistent basal meristem.

Leaf anatomy. Stomata anomocytic.

Adaxial hypodermis present. Lamina dorsiventral; without secretory cavities. Cystoliths present. The mesophyll with spherical etherial oil cells.

Stem anatomy. Secretory cavities absent. Nodes unilacunar (with two traces). Primary vascular tissue in a cylinder, without separate bundles. Secondary thickening developing from a conventional cambial ring. Xylem with fibre tracheids. Wood parenchyma paratracheal. Sieve-tube plastids P-type; type I (b).

Reproductive type, pollination. Plants andromonoecious, or polygamomonoecious.

Inflorescence, floral, fruit and seed morphology. Flowers aggregated in ‘inflorescences’; in cymes, in corymbs, and in panicles. The ultimate inflorescence unit cymose. Inflorescences dense thyrses. Flowers cyclic. Free hypanthium absent.

Perianth sepaline; (4–)8 (male flowers), or 2 (bisexual flowers); joined; 1 whorled. Calyx (if the perianth be so interpreted) (4–)8 (male flowers), or 2 (or at least, 2-lobed, in bisexual flowers); gamosepalous; persistent; in bisexual flowers accrescent (and winged); imbricate (in male flowers).

Androecium 3–5, or 6–9 (when staminodes present). Androecial members free of the perianth; free of one another; 1 whorled, or 2 whorled. Androecium exclusively of fertile stamens, or including staminodes (these sometimes present in male as well as in bisexual flowers). Staminodes when present, 3–7. Stamens 3–5. Anthers basifixed; dehiscing by longitudinal valves; latrorse. Pollen grains nonaperturate; 2-celled (in both genera).

Gynoecium 1 carpelled. The pistil 1 celled. Gynoecium monomerous; of one carpel; inferior. Carpel 1 ovuled. Placentation marginal.

Fruit non-fleshy. The fruiting carpel indehiscent (winged via the persistent, accrescent lobes of the perianth). Seeds non-endospermic. Cotyledons 2 (leafy, convolute).

Physiology, biochemistry. Alkaloids present. Aluminium accumulation not found.

Geography, cytology. Sub-tropical to tropical. Pantropical and subtropical.

Taxonomy. Subclass Dicotyledonae; Crassinucelli. Dahlgren’s Superorder Magnoliiflorae; Laurales. Cronquist’s Subclass Magnoliidae; Laurales. APG 3 core angiosperms; Superorder Magnolianae; Order unassigned to Order (formerly regarded as a synonym of Hernandiaceae, i.e. Laurales).

Species 22. Genera 2; Gyrocarpus, Sparattanthelium.

Illustrations. • Technical details: Gyrocarpus, Sparattanthelium.


This description is offered for casual browsing only. We strongly advise against extracting comparative information from it. This is much more easily achieved using the interactive key, which allows access to the character list, illustrations, full and partial descriptions, diagnostic descriptions, differences and similarities between taxa, lists of taxa exhibiting specified attributes, summaries of attributes within groups of taxa, geographical distribution, genera included in each family, classifications (Dahlgren; Dahlgren, Clifford, and Yeo; Cronquist; APG), and notes on the APG classification.

Cite this publication as: ‘Watson, L., and Dallwitz, M.J. 1992 onwards. The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Version: 25th November 2009. http://delta-intkey.com’.

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