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Anastrepha and Toxotrypana:
descriptions, illustrations, and interactive keys


Allen L. Norrbom, Cheslavo A. Korytkowski, Roberto A. Zucchi, Keiko Uramoto, George L. Venable, Jerrett McCormick and Michael J. Dallwitz

Anastrepha punensis Tigrero & Salas


Body. Setae orange brown, or red brown.

Head. Frons with brown band or mark including ocellar tubercle and extending to eye margin. Anterior or single orbital seta at margin of brown band, or within brown band. Occiput with pair of brown marks on medial and lateral sclerites, or with single broad brown area on medial and lateral sclerites (with single broad brown area, paler medially on medial sclerite). Frontal setae 3 (usually), or 4. Orbital setae 1. Ocellar seta weak, small or absent. Gena with brown spot below eye. Facial carina in profile concave or flat on dorsal 2/3. Face with ventral part gradually tapered laterally; without brown markings. Antenna not extended to ventral margin of face, or extended to or beyond ventral margin of face.

Thorax. Mesonotum length 2.8–3.7 mm (3.2–3.4, n=5; 2.93–3.26, Tigrero & Salas 2005). Postpronotal lobe and notopleuron entirely microtrichose. Scutum mostly or entirely microtrichose (sometimes with small triangular nonmicrotrichose area between dorsocentral line and presutural supra-alar seta). Scutellum disc entirely microtrichose. Postpronotal, presutural supra-alar, dorsocentral, intra-alar and scutellar setae well developed, subequal to or longer than scutellum length; postpronotal seta on posterior half of postpronotal lobe. Acrostichal seta well developed. Basal scutellar seta strong, longer than scutellum. Katepisternal seta weak, no larger than postocellar seta, or absent. Mesonotum orange brown, or medium brown, or dark brown; with short presutural lateral pale vitta on lateral margin of scutum, not extended onto notopleuron, or with presutural lateral pale vitta on lateral margin of scutum, extended to posterior notopleural seta (not reaching posterior corner nor lateral margin of notopleuron). Scutum presutural dorsocentral pale vitta present but separated anteriorly from pale area of postpronotal lobe; with 3 (both medial and sublateral) pale postsutural vittae; pale medial vitta with posterior end bilobed; pale sublateral postsutural vitta extended posteriorly to intra-alar seta. Scutum posteriorly without brown or orange brown markings (scutum predominantly brown). Scutum without brown vittae. Scutellum with at least basal third of sides and disk brown or orange, distinctly darker than apex. Propleuron at least partly dark orange or brown similar to darker area on anepisternum. Mesopleuron mostly dark orange to brown. Subscutellum entirely brown to dark brown. Mediotergite entirely brown to dark brown. Femora at least one femur partly to entirely brown (all femora and tibiae brown, tarsi yellow to orange). Fore femur with posterodorsal and ventral rows of well developed setae.

Wings. Wing length 6.5–7.4 mm (6.9–7.1, n=3; 6.64–7.2, Tigrero & Salas 2005). Wing pattern with basal half of S-band reduced to streak in cubital cells, costal band continuous to wing apex, proximal arm of V-band variable. Cell c mostly or entirely infuscated to subhyaline, or paler posteriorly, without distinct subapical hyaline area. C-band not extending to vein M in cell br adjacent to cell bm; not covering base of cell r2+3; yellow or orange area posterior to pterostigma small or absent, not extending beyond cell r1 nor distally beyond level of basal third of pterostigma. Cell bm entirely hyaline or infuscated only along subapical fold. S-band base without extension in cell a1 to or almost to posterior margin. Cubital streak (isolated base of S-band) covering at most anterior 2/3 of cell bcu, posterior third or more of bcu hyaline or faintly infuscated. Cubital streak (isolated base of S-band) extending at most slightly beyond bm-cu. Subapical hyaline area in radial cells distal to r-m extending anteriorly to vein R2+3. S-band distal section without marginal hyaline band or spots in cell r2+3 or near apices of R2+3 or R4+5. S-band distally extended to apex of vein M, or not extended to apex of vein M. V-band proximal arm as dark as apical half of S-band (extended only to or slightly anterior to vein M); extending less than 1/3 distance from apex of vein Cu1 to apex of vein A1+Cu2; not connected anteriorly to S-band; not connected to S-band in cell dm. V-band distal arm absent. S-band distal section width ratio (width of S-band/width of cell r2+3, both measured perpendicular to costal margin at apex of vein R2+3) 0.45–0.65 (0.53–0.57, n=2). Area surrounding apex of lobe of cell bcu with microtrichia similar in density to area anterdistal to it along vein Cu1. Area between S-band and V-band entirely microtrichose in cells dm and cu1. Cell c: pterostigma ratio (cell c length/pterostigma length) 1.12–1.40, n=2. Pterostigma ratio 4.2–5.4, n=2. Vein R2+3 not sinuous; without accessory vein. Vein M ratio (distance from bm-cu to r-m/distance from bm-cu to dm-cu) 0.54–0.6 (0.57–0.59, n=2). Vein M curvature ratio (width of cell r4+5 at apex/width at level of dm-cu) 0.68–0.75, n=2. Crossvein dm-cu orientation with anterior end more distal than posterior end.

Abdomen. Lateral surstylus in posterior view very short, barely extended beyond prensisetae, rounded apically. Lateral surstylus in posterior view not boot-shaped. Lateral surstylus in lateral view short, apex blunt. Phallus length 0.1–0.3 mm. Glans absent. Proctiger lateral and ventral sclerotized areas connected, lateral areas separate dorsally. Oviscape straight; length 3.6–4.2 mm (3.7–4.1, n=5; 3.89–4.18, Tigrero & Salas 2005); length ratio (oviscape length/mesonotum length) 1.05–1.35 (1.12–1.25, n=5). Eversible membrane with dorsobasal denticles mostly small and weak, apical row very large, stout, strongly sclerotized, and divided medially. Aculeus length 3.1–3.5 mm (3.35, n=2; 3.12–3.20 (tips probably broken), Tigrero & Salas 2005). Aculeus length/oviscape length 0.84, n=2. Aculeus in ventral view more or less parallel-sided except extreme base. Aculeus in lateral view straight or ventrally curved. Aculeus tip length 0.17–0.22 mm (0.19–0.20, n=2); width 0.01–0.02 mm (0.015, n=2); length/width in ventral view 12–14 (12.67–13.33, n=2); lateral margins not curved dorsally; slender, needle-like, with circular cross-section; not flared outward at or proximal to base; without ridges or lobes; without elongate dorsolateral depressions apically; not serrate; serrated part 0 times length of tip.


Sex of recorded specimens: male and female. Species group: daciformis group.

Biology and economic significance

This species is not considered economically important. Its host plants are unknown, although it probably attacks fruits of species of Sapotaceae like other species of the daciformis species group. Refer to the Fruit Fly Databases for host plant information.


• Habitus, female (dorsal). • Habitus, female (lateral). • Wing. • Terminalia, female.


Fruit Fly Databases for host plant, distribution, and nomenclatural information. Google search.